• Title: Associate Professor
  • Department: Research Institute of Ecology
  • Address: 724 Yifu Biology Building
  • Tel: 0931-8912922
  • Fax: 0931-8912922
  • Email:
  • Homepage:

2011.07 - present: Lanzhou University

Academic Appointments

2005.09-2011.06   Lanzhou University, Ecology, Ph.D.

2009.09-2011.04   University of Kentucky, Visiting Ph.D.

2001.09-2005.06   Lanzhou University, Ecology, B.S.

Administrative Appointments

Honors & Awards


The interspecies variation of seed germination timing and its effect on community structure: a case study in alpine meadow, National Natural Science Foundation of China, 630,000, 2017.1-2020.12 (Preside Over Impresario)


Effect of climate change on seed germination and seedling establishment of alpine meadow in the Qinghai-Tibetan plateau, National Natural Science Foundation of China, 370,000, 2018.1-2021.12 (Principal Participant)


Liu K.*#, Liang T#,Qiang W., Du G., Baskin J., Baskin C., Bu H., Yang H., Xiao S. (2020) Changes in seed germination strategy along the successional gradient from abandoned cropland to climax grassland in a subalpine meadow and some implications for rangeland restoration. Agriculture, Ecosystems and Environment, 289:106746

Liu, K*. , Cao, S. , Du, G. , Baskin, J. M. , Baskin, C. C. , & Bu, H. , et al. (2018). Linking seed germination and plant height: a case study of a wetland community on the eastern tibet plateau. Plant Biology, 20: 886-893.

Cao S, Liu K*, Du G, Baskin JM, Baskin CC, Bu H, Qi W (2018). Seedling emergence of 144 subalpine meadow plants: effects of phylogeny, life cycle type and seed mass. Seed Science Research 28, 93–99.

Liu, K., Baskin, J.M., Baskin, C.C., Bu H.Y., Du G.Z*, Ma M.J. Effect of Diurnal Fluctuating versus Constant Temperatures on Germination of 445 Species from the Eastern Tibet Plateau. PLoS ONE  8(7): e69364, 2013

Liu, K., Baskin, J.M., Baskin, C.C., Du G.Z*. Very fast-germinating seeds of desert species are cryptoviparous-like. Seed Science Research, 23:163-167, 2013

Liu, K., Baskin, J.M., Baskin, C.C., Bu H.Y., Liu M.X. Liu W., Du G.Z*. Effect of storage conditions on germination of 489 species from high elevation grassland of the eastern Tibet Plateau and implications for climate change . American Journal of Botany, 98: 12-19, 2011

Hai-Yan Bu*, Peng Jia, Wei Qi, Kun Liu, Dang-Hui Xu, Wen-Jing Ge, Xue-Jing Wang. The effects of phylogeny, life-history traits and altitude on the carbon, nitrogen and phosphorus contents of seeds across 203 species from an alpine meadow. Plant Ecology, 2018, 219:737-748.

Hai-Yan Bu*, Yan-Ming Zhang, Di Zhao, Su-Yu Wang, Peng Jia, Wei Qi, Kun Liu, Dang-Hui Xu, Wen-Jing Ge, Xue-Jing Wang. The evolutionary correlation associated with seed mass and altitude on nutrients allocation of seeds. Seed science research, 2018, accepted.

Hui, Z. , Robert, J. , Shidan, Z. , Hui, L. , Qiuyuan, X. ,  Wei, Q. , Kun L., Han Y. H. C, Qing Y. (2018). Shifts in functional trait–species abundance relationships over secondary subalpine meadow succession in the qinghai–tibetan plateau. Oecologia, 188:547–557.

Hui, Z. , Wei, Q. , & Kun, L. . (2018). Functional traits associated with plant colonizing and competitive ability influence species abundance during secondary succession: evidence from subalpine meadows of the qinghai-tibetan plateau. Ecology and Evolution, 8(13): 6529–6536.

Bu, H. Y. , Wang, X. J. , Zhou, X. H. , Qi, W. , Liu, K. , & Ge, W. J. , et al. (2016). The ecological and evolutionary significance of seed shape and volume for the germination of 383 species on the eastern qinghai-tibet plateau. Folia Geobotanica, 51(4), 333-341.

Haiyan Bu, Ge Wenjing, Xianhui Zhou, Wei Qi, Kun Liu, Danghui Xu, Xuejing Wang, Guozhen Du. The effect of light and seed mass on seed germination of common herbaceous species from the eastern Qinghai‐Tibet Plateau." Plant Species Biology 32.4(2016).

Zhang, Chunhui(#),Willis, Charles G.,Burghardt, Liana T.,Qi, Wei,Liu, Kun,de Moura Souza-Filho, Paulo Roberto,Ma, Zhen,Du, Guozhen*,The community-level effect of light on germination timing in relation to seed mass: a source of regeneration niche differentiation,New Phytologist,2014, 204(3):496-506.

Zhang Chunhui,Liu Kun,Qi Wei,Ma Zhen, Du Guozhen*,Light-dependent associations of germination timing with subsequent life-history traits and maternal habitats for 476 angiosperm species of the eastern Tibetan Plateau grasslands,Seed Science Research,2014, 24(3):207-215.

Qi, Wei*,Guo, Shuqing,Chen, Xuelin,Cornelissen, Johannes H. C.,Bu, Haiyan,Du, Guozhen,Cui, Xianliang,Li, Wenjin,Liu, Kun,Disentangling ecological, allometric and evolutionary determinants of the relationship between seed mass and elevation: insights from multiple analyses of 1355 angiosperm species on the eastern Tibetan Plateau,Oikos,2014, 123(1):23-32.

Wei Qi,Haiyan Bu,Kun Liu,Wenjin Li,ohannes M.H.K.,,Juhong Wang,Wenlong Li,Guozhen Du*, Biological traits are correlated with elevational distribution range of eastern Tibetan herbaceous species,Plant Ecology, 2014, 215 (10): 1187-1198.

Ma Miaojun,Zhou Xianhui,Qi Wei,Liu Kun,Jia Peng,Du Guozhen*,Seasonal Dynamics of the Plant Community and Soil Seed Bank along a Successional Gradient in a Subalpine Meadow on the Tibetan Plateau,PLos One,2013, 8(11).

Bu H.Y., Du, G.Z*., Chen, X.L.,  Wang, Y.F., Xu, X.L.,  Liu, K. The Evolutionary Significance of Seed Germinability in an Alpine Meadow on the eastern Qinghai-Tibet Plateau. Arctic, Antarctic, and Alpine Research, 41 : 97–102, 2009

Haiyan Bu,XuelinChen,XiuliXu,Kun Liu,Peng Jia Guo,zhen Du. Seed mass and germination in an alpine meadow on the eastern Tsinghai–Tibet plateau. Plant Ecology 191:127–149, 2007

Haiyan Bu,Guozhen Du,Xuelin Chen,Xiuli Xu,Kun Liu,Shujun Wen. Community-wide germination strategies in an alpine meadow on the eastern Qinghai-Tibet plateau: phylogenetic and life-history correlates. Plant Ecology 195:87-98, 2008

H. Y. Bu, X. L. Chen, Y. Wang, X. L. Xu, K. Liu and G. Z. Du. Germination time, other plant traits and phylogeny in an alpine meadow on the eastern Qinghai-Tibet plateau. Community Ecology 8(2):221-227, 2007

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